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Forages on ground and in trees and shrubs (Terres 1980).","animalPhenologyComments":null,"colonialBreeder":false,"length":22,"width":null,"weight":45,"animalPhenologies":[{"animalCagPhenologyId":102135,"animalPhenology":{"id":3,"animalPhenologyDescEn":"Diurnal","animalPhenologyDescEs":"Diurno","animalPhenologyDescFr":"Diurne","displayOrder":3},"adult":true,"immature":true}],"animalFoodHabits":[{"animalCagFoodHabitsId":103965,"foodHabits":{"id":3,"foodHabitsDescEn":"Invertivore","foodHabitsDescEs":"Invertívoro","foodHabitsDescFr":"Invertivore","displayOrder":3},"adult":true,"immature":true},{"animalCagFoodHabitsId":103966,"foodHabits":{"id":5,"foodHabitsDescEn":"Granivore","foodHabitsDescEs":"Granívoro","foodHabitsDescFr":"Granivore","displayOrder":5},"adult":true,"immature":true},{"animalCagFoodHabitsId":103967,"foodHabits":{"id":6,"foodHabitsDescEn":"Frugivore","foodHabitsDescEs":"Frugívoro","foodHabitsDescFr":"Frugivore","displayOrder":6},"adult":true,"immature":true}]},"occurrenceDelineations":[{"eoSpecsDetailId":124587,"locationUseClass":{"id":2,"locationUseClassDescEn":"Breeding","locationUseClassDescEs":"En reproducción","locationUseClassDescFr":"Reproduction","displayOrder":2},"eoSpecGroupName":"Passerines","subtypes":"Foraging Area, Nest Site, Nesting Colony","inferredExtentDistance":null,"inferredExtentNotes":null,"minimumEoCriteria":"Evidence of historical breeding, or current and likely recurring breeding, at a given location, minimally a reliable observation of one or more breeding pairs in appropriate habitat. Be cautious about creating EOs for observations that may represent single breeding events outside the normal breeding distribution.","mappingGuidance":"Breeding occurrences include nesting areas as well as foraging areas.<br/><br/>For swallows and other species that have separate nesting and foraging areas, separations are based on nest sites or nesting areas, not to locations of foraging individuals. For example, nesting areas separated by a gap larger than the separation distance are different occurrences, regardless of the foraging locations of individuals from those nesting areas. This separation procedure is appropriate because nesting areas are the critical aspect of swallow breeding occurrences, tend to be relatively stable or at least somewhat predictable in general location, and so are the basis for effective conservation; foraging areas are much more flexible and not necessarily static.","separationBarriers":null,"separationDistanceUnsuitableHabitatat":5.0,"separationDistanceSuitableHabitatat":5.0,"altSeparationProcedure":null,"separationJustification":"Significant dispersal and associated high potential for gene flow among populations of birds separated by tens of kilometers (e.g., Moore and Dolbeer 1989), and increasing evidence that individuals leave their usual home range to engage in extrapair copulations, as well as long foraging excursions of some species, make it difficult to circumscribe occurrences on the basis of meaningful population units without occurrences becoming too large. Hence, a moderate, standardized separation distance has been adopted for songbirds and flycatchers; it should yield occurrences that are not too spatially expansive while also accounting for the likelihood of gene flow among populations within a few kilometers of each other.<br/><br/>Be careful not to separate a population's nesting areas and foraging areas as different occurrences; include them in the same occurrence even if they are more than 5 km apart. Mean foraging radius (from nesting area) of Brown-headed Cowbird females was 4.0 kilometers in California, 1.2 kilometers in Illinois-Missouri (Thompson 1994). Yellow-headed Blackbirds, Brewer's Blackbirds, and probably Red-winged Blackbirds all forage up to 1.6 kilometers away from breeding colony (Willson 1966, Horn 1968). In one study, Brewer's Blackbirds were found as far as 10 kilometers from nesting area (Williams 1952), but this may be unusual.<br/><br/>For swallows and other passerines with similar behavioral ecology, separation distance pertains to nest sites or nesting colonies, not to locations of foraging individuals. For example, nesting areas separated by a gap of more than 5 km are different occurrences, regardless of the foraging locations of individuals from those nesting areas. This separation procedure is appropriate because nesting areas are the critical aspect of swallow breeding occurrences, tend to be relatively stable or at least somewhat predictable in general location, and so are the basis for effective conservation; foraging areas are much more flexible and not necessarily static.<br/><br/>Be cautious about creating EOs for observations that may represent single breeding events outside the normal breeding distribution.<br/><br/>Unsuitable habitat: Habitat not normally used for breeding/feeding by a particular species. For example, unsuitable habitat for grassland and shrubland birds includes forest/woodland, urban/suburban, and aquatic habitats. Most habitats would be suitable for birds with versatile foraging habits (e.g., most corvids).","versionDate":"2004-09-10","versionAuthor":"Hammerson, G.","versionNotes":null,"lastModified":"2025-12-05T22:48:36.225047Z"},{"eoSpecsDetailId":124588,"locationUseClass":{"id":3,"locationUseClassDescEn":"Nonbreeding","locationUseClassDescEs":"No reproducción","locationUseClassDescFr":"Non-reproduction","displayOrder":3},"eoSpecGroupName":"Passerines","subtypes":"Foraging Area, Roost Site","inferredExtentDistance":null,"inferredExtentNotes":null,"minimumEoCriteria":"Any area used traditionally in the nonbreeding season (used for populations that are not resident in a location year-round). Minimally, reliable observations of 10 or more individuals in appropriate habitat for 20 or more days at a time. For G1-G3 species, observations of fewer individuals could constitute an occurrence of conservation value. Sites used during migration should be documented under the 'migratory stopover' location use class.","mappingGuidance":null,"separationBarriers":null,"separationDistanceUnsuitableHabitatat":5.0,"separationDistanceSuitableHabitatat":5.0,"altSeparationProcedure":null,"separationJustification":"Separation distance is necessarily arbitrary but attempts to balance the high mobility of birds with the need for occurrences of reasonable spatial scope. Note that a population's roost sites and foraging areas are parts of the same occurrence, even if they are more than 5 km apart.<br/><br/>For swallows and other species with similar behavioral ecology, the separation distance pertains to communal roost sites rather than to foraging areas; the former tend to be more stable and specific over time than the latter.","versionDate":"2004-09-03","versionAuthor":"Hammerson, G.","versionNotes":null,"lastModified":"2025-12-05T22:48:36.225047Z"},{"eoSpecsDetailId":124589,"locationUseClass":{"id":4,"locationUseClassDescEn":"Migratory stopover","locationUseClassDescEs":"Parada migratoria","locationUseClassDescFr":"Halte migratoire","displayOrder":5},"eoSpecGroupName":"Passerines","subtypes":"Foraging Area, Roost Site","inferredExtentDistance":null,"inferredExtentNotes":null,"minimumEoCriteria":"For most passerines: Evidence of recurring presence of migrating individuals (including historical) and potential recurring presence at a given location; minimally a reliable observation of 25 birds in appropriate habitat.<br/><br/>For swallows: Evidence of recurring presence of migrating flocks (including historical) and potential recurring presence at a given location; minimally a reliable observation of 100 birds in appropriate habitat (e.g., traditional roost sites).<br/><br/>Occurrences should be locations where the species is resident for some time during the appropriate season; it is preferable to have observations documenting presence over at least 7 days annually.<br/><br/>EOs should not be described for species that are nomadic during nonbreeding season: e.g., Lark Bunting.<br/><br/>Be cautious about creating EOs for observations that may represent single events.","mappingGuidance":null,"separationBarriers":null,"separationDistanceUnsuitableHabitatat":5.0,"separationDistanceSuitableHabitatat":5.0,"altSeparationProcedure":null,"separationJustification":"Separation distance somewhat arbitrary but intended to define occurrences of manageable size for conservation purposes. Occurrences defined primarily on the basis of areas supporting concentrations of birds, rather than on the basis of distinct populations.<br/><br/>For swallows and other species with similar behavioral ecology, the separation distance pertains to communal roost sites rather than to foraging areas; the former tend to be more stable and specific over time than the latter.","versionDate":"2004-09-03","versionAuthor":"Hammerson, G., and S. Cannings","versionNotes":null,"lastModified":"2025-12-05T22:48:36.225047Z"},{"eoSpecsDetailId":124586,"locationUseClass":{"id":13,"locationUseClassDescEn":"Nonmigratory","locationUseClassDescEs":"No migratorio","locationUseClassDescFr":"Non migratrice","displayOrder":20},"eoSpecGroupName":"Passerines","subtypes":null,"inferredExtentDistance":null,"inferredExtentNotes":null,"minimumEoCriteria":"Occurrences are based on evidence of historical presence, or current and likely recurring presence, at a particular location. Such evidence minimally includes collection or reliable observation and documentation of one or more individuals in or near appropriate habitat.<br/><br/>These occurrence specifications are used for nonmigratory populations of passerine birds.","mappingGuidance":null,"separationBarriers":"None.","separationDistanceUnsuitableHabitatat":5.0,"separationDistanceSuitableHabitatat":5.0,"altSeparationProcedure":null,"separationJustification":"Significant dispersal and associated high potential for gene flow among populations of birds separated by tens of kilometers (e.g., Moore and Dolbeer 1989), and increasing evidence that individuals leave their usual home range to engage in extrapair copulations, as well as long foraging excursions of some species, make it difficult to circumscribe occurrences on the basis of meaningful population units without occurrences becoming too large. Hence, a moderate, standardized separation distance has been adopted for songbirds and flycatchers; it should yield occurrences that are not too spatially expansive while also accounting for the likelihood of gene flow among populations within a few kilometers of each other.<br/><br/>Be careful not to separate a population's nesting areas and breeding-season foraging areas as different occurrences; include them in the same occurrence even if they are more than 5 km apart. Blue jays have small summer home ranges but fly up to 4 kilometers to harvest mast (Tarvin and Woolfenden 1999). Flocks of pinyon jays range over 21-29 square kilometers (Ligon 1971, Balda and Bateman 1971); nesting and foraging areas may be widely separated. Tricolored blackbirds forage in flocks that range widely to more than 15 kilometers from the nesting colony (Beedy and Hamilton 1999).<br/><br/>Unsuitable habitat: Habitat not normally used for breeding/feeding by a particular species. For example, unsuitable habitat for grassland and shrubland birds includes forest/woodland, urban/suburban, and aquatic habitats. Most habitats would be suitable for birds with versatile foraging habits (e.g., most corvids).","versionDate":"2004-09-10","versionAuthor":"Hammerson, G.","versionNotes":"These specs pertain to nonmigratory species.","lastModified":"2025-12-05T22:48:36.225047Z"}],"plantCharacteristics":null,"elementManagement":{"elementGlobalId":105565,"eoManagementGroupName":null,"stewardshipOverview":null,"impacts":null,"restorationPotential":null,"siteConservationPlansConsidered":null,"managementMethods":null,"monitoringMethods":null,"managementProgramContacts":null,"monitoringProgramContacts":null,"managementResearchPrograms":null,"managementResearchNeeds":null,"biologicalResearchNeeds":null,"additionalTopics":null},"occurrenceViabilities":[{"eoRankSpecsDetailId":106987,"eoRankSpecsGroupName":null,"locationUseClass":{"id":1,"locationUseClassDescEn":"Not applicable","locationUseClassDescEs":"No se aplica","locationUseClassDescFr":"Sans objet","displayOrder":1},"excellentViability":null,"goodViability":null,"fairViability":null,"poorViability":null,"viabilityJustification":"Use the Generic Guidelines for the Application of Occurrence Ranks (2008).<br>The Key for Ranking Species Occurrences Using the Generic Approach provides a step-wise process for implementing this method.","versionDate":null,"versionAuthor":null,"versionNotes":null,"lastModified":"2020-05-14T02:29:18.481339Z"}],"references":[{"id":153779,"citation":"American Ornithologists' Union (AOU). 1983. 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Condor 73:287-302.","shortCitationAuthor":"Balda and Bateman","shortCitationYear":1971,"referenceCode":"A71BAL01NACA","lastModified":"2020-02-25T14:38:12.657866Z"},{"id":137951,"citation":"Banks, R. C., and M. R. Browning. 1995. Comments on the status of revived old names for some North American birds. Auk 112:633-648.","shortCitationAuthor":"Banks and Browning","shortCitationYear":1995,"referenceCode":"A95BAN01NAUS","lastModified":"2020-02-25T14:38:12.673494Z"},{"id":153590,"citation":"Bent, A.C., et al. 1968. Life histories of North American cardinals, grosbeaks, buntings, towhees, finches, sparrows, and allies. Part Two. U.S. National Museum Bulletin 237. (reprinted by Dover Publications, Inc., New York, NY).","shortCitationAuthor":"Bent et al.","shortCitationYear":1968,"referenceCode":"B68BEN01NAUS","lastModified":"2020-02-25T18:12:35.976174Z"},{"id":463505,"citation":"BirdLife International. 2004b. Threatened birds of the world 2004. CD ROM. BirdLife International, Cambridge, UK.","shortCitationAuthor":"BirdLife International","shortCitationYear":2004,"referenceCode":"N04BIR0100LA","lastModified":"2020-02-25T14:38:12.704742Z"},{"id":150327,"citation":"Dow, D. D. 1969. Home range and habitat of the Cardinal in peripheral and central populations. Canadian Journal of Zoology 47:103-114.","shortCitationAuthor":"Dow","shortCitationYear":1969,"referenceCode":"A69DOW01NACA","lastModified":"2020-02-25T18:25:14.206563Z"},{"id":102357,"citation":"Filliater, T. S., R. Breitwisch, and P. M. Nealen. 1994. Predation on northern cardinal nests: does choice of nest site matter? Condor 96:761-768.","shortCitationAuthor":"Filliater et al.","shortCitationYear":1994,"referenceCode":"A94FIL01NAUS","lastModified":"2020-02-25T18:25:14.206563Z"},{"id":100210,"citation":"Harrison, C. 1978. A Field Guide to the Nests, Eggs and Nestlings of North American Birds. Collins, Cleveland, Ohio.","shortCitationAuthor":"Harrison","shortCitationYear":1978,"referenceCode":"B78HAR01NAUS","lastModified":"2020-02-25T14:38:12.657866Z"},{"id":107035,"citation":"Harrison, H. H. 1979. A field guide to western birds' nests. Houghton Mifflin Company, Boston. 279 pp.","shortCitationAuthor":"Harrison","shortCitationYear":1979,"referenceCode":"B79HAR01NAUS","lastModified":"2020-02-25T18:11:56.303852Z"},{"id":105464,"citation":"Horn, H. S. 1968. The adaptive significance of colonial nesting in the Brewer's Blackbird. Ecology 49:682-694.","shortCitationAuthor":"Horn","shortCitationYear":1968,"referenceCode":"A68HOR01NACA","lastModified":"2020-02-25T14:38:12.689104Z"},{"id":155592,"citation":"Howell, S. N. G., and S. Webb. 1995.  A guide to the birds of Mexico and northern Central America. 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In Ohio, nesting success rate was only 15% (Filliater et al. 1994). <br><br>NON-BREEDING: Winter home ranges in Kentucky averaged 21.2 hectares, although these may have been underestimated because of limited observation time (Ritchison and Omer 1990).","habitatComments":"Thickets, brushy areas, fields, shrubbery, forest edge, clearings, around human habitation, and, in arid regions, in scrub, riparian thickets, woodland, and brush; typically avoids dense forest (but penetrates native forest in Hawaii). <br><br>Nests in dense shrubs, small deciduous or coniferous tree, thicket, vine, briar tangle, mesquite, generally 3 m or less above ground. 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Check-list of North American birds. Seventh edition. American Ornithologists' Union, Washington, D.C. [as modified by subsequent supplements and corrections published in <i>The Auk</i>]. Also available online: http://www.aou.org/.","shortCitationAuthor":"American Ornithologists' Union (AOU)","shortCitationYear":1998,"referenceCode":"B98AOU01NAUS","lastModified":"2020-02-25T14:38:12.673494Z"},"predecessors":[],"elementGlobalThreatAssessments":[]}