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Louisiana, and Mississippi, and through eastern Mexico to the Isthmus of Tehuantepec, then discontinuously through southern Mexico, Belize, Guatemala, El Salvador, Honduras, and Nicaragua to Costa Rica, at elevations from sea level to 2,438 meters (8,000 feet) (Rossman et al. 1996).","areaOfOccupancy":null,"areaOfOccupancyComments":null,"numberEosComments":"This species is represented by a very large number of occurrences (Rossman 1970). Werler and Dixon (2000) mapped hundreds of collection sites in Texas, and Trauth et al. (2004) mapped well over 100 collection sites in Arkansas.","popSizeComments":"Total adult population size is unknown but certainly exceeds 100,000. This snake is common in the southern United States.","viabilityComments":null,"threatImpactComments":"No major threats are known. Locally, wetland loss and degradation have extirpated or reduced some populations.","shortTermTrendComments":"Currently, the extent of occurrence, area of occupancy, number of subpopulations, and population size probably are relatively stable or declining at a rate of less than 10 percent over 10 years or three generations.","longTermTrendComments":"Area of occupancy, number of subpopulations, and population size have declined modestly, especially at the margins of the range. For example, this species declined at the northern fringe of the range in northwestern Indiana between the 1930s and 1990s (Brodman et al. 2002).","inventoryNeeds":null,"numberProtEosComments":"Many occurrences are in protected areas.","protectionNeeds":null,"otherConsiderations":null,"intrinsicVulnerabilityComments":null,"enviromentalSpecificityComments":null},"animalCharacteristics":{"elementGlobalId":104971,"majorHabitat":{"id":3,"majorHabitatDescEn":"Terrestrial","majorHabitatDescEs":"Terrestre","majorHabitatDescFr":"Terrestre"},"nonMigrant":false,"localMigrant":true,"longDistanceMigrant":false,"mobilityMigrationComments":"May migrate between separate winter and summer habitats (Vogt 1981).","foodHabitsComments":"Eats frogs, toads, tadpoles, salamanders, fishes, insects, earthworms, occasionally carrion (Stebbins 1985, Collins 1982).","animalPhenologyComments":"Active day or night; nocturnal in hot weather (Collins 1982). Active from March to October in north (Collins 1982).","colonialBreeder":false,"length":123,"width":null,"weight":null,"animalPhenologies":[{"animalCagPhenologyId":103181,"animalPhenology":{"id":1,"animalPhenologyDescEn":"Hibernates/aestivates","animalPhenologyDescEs":"Hibernación/Estivación","animalPhenologyDescFr":"Hiberne/estive","displayOrder":1},"adult":true,"immature":true},{"animalCagPhenologyId":103182,"animalPhenology":{"id":3,"animalPhenologyDescEn":"Diurnal","animalPhenologyDescEs":"Diurno","animalPhenologyDescFr":"Diurne","displayOrder":3},"adult":true,"immature":true},{"animalCagPhenologyId":103183,"animalPhenology":{"id":4,"animalPhenologyDescEn":"Nocturnal","animalPhenologyDescEs":"Nocturno","animalPhenologyDescFr":"Nocturne","displayOrder":4},"adult":true,"immature":true},{"animalCagPhenologyId":103184,"animalPhenology":{"id":5,"animalPhenologyDescEn":"Crepuscular","animalPhenologyDescEs":"Crepuscular","animalPhenologyDescFr":"Crépusculaire","displayOrder":5},"adult":true,"immature":true}],"animalFoodHabits":[{"animalCagFoodHabitsId":105888,"foodHabits":{"id":1,"foodHabitsDescEn":"Carnivore","foodHabitsDescEs":"Carnívoro","foodHabitsDescFr":"Carnivore","displayOrder":1},"adult":true,"immature":true},{"animalCagFoodHabitsId":105889,"foodHabits":{"id":2,"foodHabitsDescEn":"Piscivore","foodHabitsDescEs":"Piscivore","foodHabitsDescFr":"Piscivore","displayOrder":2},"adult":true,"immature":true},{"animalCagFoodHabitsId":105890,"foodHabits":{"id":3,"foodHabitsDescEn":"Invertivore","foodHabitsDescEs":"Invertívoro","foodHabitsDescFr":"Invertivore","displayOrder":3},"adult":true,"immature":true}]},"occurrenceDelineations":[{"eoSpecsDetailId":1142,"locationUseClass":{"id":1,"locationUseClassDescEn":"Not applicable","locationUseClassDescEs":"No se aplica","locationUseClassDescFr":"Sans objet","displayOrder":1},"eoSpecGroupName":"Medium And Large Colubrid Snakes","subtypes":null,"inferredExtentDistance":0.5,"inferredExtentNotes":null,"minimumEoCriteria":"Occurrences are based on evidence of historical presence, or current and likely recurring presence, at a given location. Such evidence minimally includes collection or reliable observation and documentation of one or more individuals (including eggs) in or near appropriate habitat where the species is presumed to be established and  breeding.","mappingGuidance":null,"separationBarriers":"Busy highway or highway with obstructions such that snakes rarely if ever cross successfully; major river, lake, pond, or deep marsh (this barrier pertains only to upland species and does not apply to aquatic or wetland snakes); densely urbanized area dominated by buildings and pavement.","separationDistanceUnsuitableHabitatat":1.0,"separationDistanceSuitableHabitatat":10.0,"altSeparationProcedure":null,"separationJustification":"Available information on movements of colubrid snakes is limited to a small minority of species. These data indicate that nearly all species have home ranges smaller or much smaller than 25 ha (e.g., less than 3 ha, <i>Pituophis catenifer</i> in California, Rodriguez-Robles 2003), with some up to about 75 ha (<i>Heterodon platirhinos</i>, average 50 ha, Plummer and Mills 2000), and the largest up to 225 ha in the biggest colubrids (<i>Drymarchon</i>, summer mean 50-100 ha, USFWS 1998). <br><br>Radiotelemetry data for <i>Pantherophis </i>indicate that residents of hibernacula that are 1-2 km apart (with suitable intervening habitat) probably interbreed (Prior et al. 1997, Blouin-Demers and Weatherhead 2002). However, \"evidence of genetic structure even over short distances (e.g., 2-20 km) implies that gene flow among rat snake populations can be easily disrupted\" (Prior et al. 1997). Loughheed et al. (1999) found evidence of substantial genetic exchange among local hibernacula (< 6 km apart), but gene flow over distances of 10s of km appears to be substantially less. Based on extensive radio-tracking data, Blouin-Demers and Weatherhead (2002) found that home range size of <i>Pantherophis </i>averaged 18.5 ha and ranged up to 93 ha; based on the most mobile individuals, <i>Pantherophis </i>from hibernacula up to 8 km apart can come together for mating. <i>Pantherophis </i>and probably other colubrids exhibit high fidelity to hibernacula and shift even to nearby sites only rarely (Prior et al. 2001). <br><br>Many of the several studies that report small home ranges for colubrids did not employ methods (e.g., radio telemetry) suitable for detecting full annual or multi-annual home range size, dispersal, or other long-distance movements, so these may have yielded underestimates of home ranges or activity areas.  <br><br>At least some colubrids, including medium-sized species such as garter snakes, not uncommonly move between areas up to a few kilometers apart, and several species make extensive movements of up to several kilometers, so separation distances of 1-2 km for suitable habitat are too small for medium-sized and large colubrids.<br><br>A separation distance of 10 km for suitable habitat was selected as most appropriate for snakes assigned to this Specs Group because it seems generally unlikely that two locations separated by less than 10 km of suitable habitat would represent distinct occurrences.<br><br>For the purposes of these occurrence specifications, upland habitat is regarded as unsuitable habitat for aquatic and wetland snakes. For upland snakes, shallow or patchy wetlands are treated as unsuitable habitat whereas large deepwater habitats (subjective determination) are barriers.","versionDate":"2013-02-12","versionAuthor":"Hammerson, G.","versionNotes":"Separation distance for suitable habitat was changed from 5 km to 10 km based on comments from Dale Jackson (12 Feb 2013).","lastModified":"2020-04-30T02:04:05.987846Z"}],"plantCharacteristics":null,"elementManagement":{"elementGlobalId":104971,"eoManagementGroupName":null,"stewardshipOverview":null,"impacts":null,"restorationPotential":null,"siteConservationPlansConsidered":null,"managementMethods":null,"monitoringMethods":null,"managementProgramContacts":null,"monitoringProgramContacts":null,"managementResearchPrograms":null,"managementResearchNeeds":null,"biologicalResearchNeeds":null,"additionalTopics":null},"occurrenceViabilities":[{"eoRankSpecsDetailId":106409,"eoRankSpecsGroupName":null,"locationUseClass":{"id":1,"locationUseClassDescEn":"Not applicable","locationUseClassDescEs":"No se aplica","locationUseClassDescFr":"Sans objet","displayOrder":1},"excellentViability":null,"goodViability":null,"fairViability":null,"poorViability":null,"viabilityJustification":"Use the Generic Guidelines for the Application of Occurrence Ranks (2008).<br>The Key for Ranking Species Occurrences Using the Generic Approach provides a step-wise process for implementing this method.","versionDate":null,"versionAuthor":null,"versionNotes":null,"lastModified":"2020-05-14T02:27:55.323641Z"}],"references":[{"id":137372,"citation":"Behler, J. 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Amphibians and reptiles in Kansas. Second edition. Univ. Kansas Mus. Nat. Hist., Pub. Ed. Ser. 8. xiii + 356 pp.","shortCitationAuthor":"Collins","shortCitationYear":1982,"referenceCode":"B82COL01NAUS","lastModified":"2020-02-25T18:12:09.428867Z"},{"id":137880,"citation":"Collins, J. T. 1990. Standard common and current scientific names for North American amphibians and reptiles. 3rd ed. Society for the Study of Amphibians and Reptiles. Herpetological Circular No. 19. 41 pp.","shortCitationAuthor":"Collins","shortCitationYear":1990,"referenceCode":"B90COL01NAUS","lastModified":"2020-02-25T14:38:15.579798Z"},{"id":140525,"citation":"Conant, R., and J. T. Collins. 1998. A field guide to reptiles and amphibians: eastern and central North America. Third edition, expanded. 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