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Toronto. 1150 pp.","shortCitationAuthor":"Lindzey","shortCitationYear":1987,"referenceCode":"A87LIN61HQUS","lastModified":"2020-02-25T18:15:48.276145Z"},{"id":810393,"citation":"Linzey, D.W. 2016. Mammals of Great Smoky Mountains National Park: 2016 revision. Southeastern Naturalist 15(Monograph 8):1–93.","shortCitationAuthor":"Linzey","shortCitationYear":2016,"referenceCode":"A16LIN02EHUS","lastModified":"2020-02-25T18:12:52.46115Z"},{"id":387023,"citation":"Logan, K. A., and L. L. Sweanor. 2001. Desert puma: evolutionary ecology and conservation of an enduring carnivore. Island Press, Covelo, California. 463 pp.","shortCitationAuthor":"Logan and Sweanor","shortCitationYear":2001,"referenceCode":"B01LOG01NAUS","lastModified":"2020-02-25T18:15:48.276145Z"},{"id":125960,"citation":"Maehr, D. S., and J. B. Wooding. 1992. Florida black bear <i>Ursus americanus floridanus</i>. Pages 265-275 in S. R. Humphrey, editor. Rare and endangered biota of Florida. Vol. I. Mammals. Univ. 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