Lepomis gibbosus - (Linnaeus, 1758)
Pumpkinseed
Taxonomic Status: Accepted
Related ITIS Name(s): Lepomis gibbosus (Linnaeus, 1758) (TSN 168144)
Unique Identifier: ELEMENT_GLOBAL.2.105048
Element Code: AFCQB11030
Informal Taxonomy: Animals, Vertebrates - Fishes - Bony Fishes - Sunfishes and Freshwater Basses
Image 86

© Noel Burkhead & Virginia Dept of Game and Inland Fisheries (Fishes of Virginia)

 
Kingdom Phylum Class Order Family Genus
Animalia Craniata Actinopterygii Perciformes Centrarchidae Lepomis
Genus Size: C - Small genus (6-20 species)
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Concept Reference
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Concept Reference: Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea, and W.B. Scott. 1991. Common and scientific names of fishes from the United States and Canada. American Fisheries Society, Special Publication 20. 183 pp.
Concept Reference Code: B91ROB01NAUS
Name Used in Concept Reference: Lepomis gibbosus
Taxonomic Comments: Though the gender of the name LEPOMIS is feminine (see Bailey and Robins, 1988, Bull. Zool. Nomencl. 45(2):100), the 1991 AFS checklist (Robins et al. 1991) retained the masculine ending for GIBBOSUS and other species, pending a vote by the ICZN on a petition (by Etnier and Warren) to treat LEPOMIS as masculine for nomenclatural purposes.
Conservation Status
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NatureServe Status

Global Status: G5
Global Status Last Reviewed: 09Feb2016
Global Status Last Changed: 23Sep1996
Ranking Methodology Used: Ranked by inspection
Rounded Global Status: G5 - Secure
Nation: United States
National Status: N5 (05Dec1996)
Nation: Canada
National Status: N5 (18Aug2015)

U.S. & Canada State/Province Status
United States Alabama (SNA), Arizona (SNA), California (SNA), Colorado (SNA), Connecticut (S5), Delaware (S5), District of Columbia (S5), Georgia (S4), Idaho (SNA), Illinois (S3S4), Indiana (S4), Iowa (SNR), Kentucky (S4), Maine (S5), Maryland (S5), Massachusetts (S5), Michigan (S5), Minnesota (SNR), Montana (SNA), Nebraska (SNR), Nevada (SNA), New Hampshire (S5), New Jersey (S5), New York (S5), North Carolina (S5), North Dakota (SNR), Ohio (S5), Oregon (SNA), Pennsylvania (S5), Rhode Island (S5), South Carolina (SNR), Tennessee (SNA), Texas (SNA), Vermont (S5), Virginia (S5), Washington (SNA), West Virginia (S5), Wisconsin (S5), Wyoming (SNA)
Canada British Columbia (SNA), Manitoba (S1), New Brunswick (S5), Ontario (S5), Quebec (S5)

Other Statuses

IUCN Red List Category: LC - Least concern

NatureServe Global Conservation Status Factors

Range Extent Comments: Native to much of Atlantic slope from New Brunswick to Edisto River, South Carolina; Great Lakes, Hudson Bay, and upper Mississippi basins from Quebec and New York to southeastern Manitoba and North Dakota, and south to northern Kentucky and Missouri drainage; common (Page and Burr 1991). Widely introduced in western U.S., southern Canada, and Europe.

Number of Occurrences: 81 to >300
Number of Occurrences Comments: This species is represented by a large number of subpopulations and locations.

Population Size: >1,000,000 individuals
Population Size Comments: Total adult population size is unknown but very large.

Overall Threat Impact Comments: Localized threats may exist, but on a range-wide scale no major threats are known.

Short-term Trend Comments: Trend over the past 10 years or three generations is uncertain but likely relatively stable or slowly declining.

Other NatureServe Conservation Status Information

Distribution
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Global Range: Native to much of Atlantic slope from New Brunswick to Edisto River, South Carolina; Great Lakes, Hudson Bay, and upper Mississippi basins from Quebec and New York to southeastern Manitoba and North Dakota, and south to northern Kentucky and Missouri drainage; common (Page and Burr 1991). Widely introduced in western U.S., southern Canada, and Europe.

U.S. States and Canadian Provinces
Color legend for Distribution Map
Endemism: occurs (regularly, as a native taxon) in multiple nations

U.S. & Canada State/Province Distribution
United States ALexotic, AZexotic, CAexotic, COexotic, CT, DC, DE, GA, IA, IDexotic, IL, IN, KY, MA, MD, ME, MI, MN, MTexotic, NC, ND, NE, NH, NJ, NVexotic, NY, OH, ORexotic, PA, RI, SC, TNexotic, TXexotic, VA, VT, WAexotic, WI, WV, WYexotic
Canada BCexotic, MB, NB, ON, QC

Range Map
No map available.

U.S. Distribution by Watershed Help
Watershed Region Help Watershed Name (Watershed Code)
01 West Branch Penobscot (01020001), East Branch Penobscot (01020002), Mattawamkeag (01020003), Piscataquis (01020004), Lower Penobscot (01020005), Upper Kennebec (01030001), Dead (01030002), Lower Kennebec (01030003), Upper Androscoggin (01040001), Lower Androscoggin (01040002), St. Croix (01050001), Maine Coastal (01050002), St. George-Sheepscot (01050003), Presumpscot (01060001), Saco (01060002), Piscataqua-Salmon Falls (01060003), Pemigewasset (01070001), Merrimack (01070002), Contoocook (01070003), Nashua (01070004), Concord (01070005), Upper Connecticut (01080101), Passumpsic (01080102), Waits (01080103), Upper Connecticut-Mascoma (01080104), White (01080105), Black-Ottauquechee (01080106), West (01080107), Middle Connecticut (01080201), Miller (01080202), Deerfield (01080203), Chicopee (01080204), Lower Connecticut (01080205), Westfield (01080206), Farmington (01080207), Charles (01090001), Cape Cod (01090002), Blackstone (01090003), Narragansett (01090004), Pawcatuck-Wood (01090005), Quinebaug (01100001), Shetucket (01100002), Thames (01100003), Quinnipiac (01100004), Housatonic (01100005), Saugatuck (01100006), St. Francois (01110000)
02 Lake George (02010001), Otter (02010002), Winooski (02010003), Ausable (02010004), Lamoille (02010005), Great Chazy-Saranac (02010006), Missisquoi (02010007), Upper Hudson (02020001), Sacandaga (02020002), Hudson-Hoosic (02020003), Mohawk (02020004), Schoharie (02020005), Middle Hudson (02020006), Rondout (02020007), Hudson-Wappinger (02020008), Lower Hudson (02030101), Bronx (02030102), Hackensack-Passaic (02030103), Raritan (02030105), Upper Delaware (02040101), East Branch Delaware (02040102), Lackawaxen (02040103), Middle Delaware-Mongaup-Brodhead (02040104), Middle Delaware-Musconetcong (02040105), Lehigh (02040106), Crosswicks-Neshaminy (02040201), Lower Delaware (02040202), Schuylkill (02040203), Brandywine-Christina (02040205), Cohansey-Maurice (02040206), Broadkill-Smyrna (02040207), Mullica-Toms (02040301), Great Egg Harbor (02040302), Upper Susquehanna (02050101), Chenango (02050102), Owego-Wappasening (02050103), Tioga (02050104), Chemung (02050105), Upper Susquehanna-Tunkhannock (02050106), Upper Susquehanna-Lackawanna (02050107), Upper West Branch Susquehanna (02050201), Sinnemahoning (02050202), Middle West Branch Susquehanna (02050203), Bald Eagle (02050204), Pine (02050205), Lower West Branch Susquehanna (02050206), Lower Susquehanna-Penns (02050301), Upper Juniata (02050302), Raystown (02050303), Lower Juniata (02050304), Lower Susquehanna-Swatara (02050305), Lower Susquehanna (02050306), Chester-Sassafras (02060002), Gunpowder-Patapsco (02060003), Severn (02060004), Choptank (02060005), Patuxent (02060006), Blackwater-Wicomico (02060007), Nanticoke (02060008), Pocomoke (02060009), Chincoteague (02060010), North Branch Potomac (02070002), Cacapon-Town (02070003), Conococheague-Opequon (02070004), South Fork Shenandoah (02070005), North Fork Shenandoah (02070006), Shenandoah (02070007), Middle Potomac-Catoctin (02070008), Monocacy (02070009), Middle Potomac-Anacostia-Occoquan (02070010), Lower Potomac (02070011), Great Wicomico-Piankatank (02080102), Rapidan-Upper Rappahannock (02080103), Lower Rappahannock (02080104), Mattaponi (02080105), Pamunkey (02080106), York (02080107), Lynnhaven-Poquoson (02080108), Western Lower Delmarva (02080109), Eastern Lower Delmarva (02080110), Upper James (02080201), Maury (02080202), Middle James-Buffalo (02080203), Rivanna (02080204), Middle James-Willis (02080205), Lower James (02080206), Appomattox (02080207), Hampton Roads (02080208)
03 Upper Roanoke (03010101), Middle Roanoke (03010102), Upper Dan (03010103), Lower Dan (03010104), Banister (03010105), Roanoke Rapids (03010106), Lower Roanoke (03010107), Nottoway (03010201), Blackwater (03010202), Ghowan (03010203), Meheriin (03010204), Albemarle (03010205), Upper Tar (03020101), Fishing (03020102), Lower Tar (03020103), Pamlico (03020104), Pamlico Sound (03020105), Bogue-Core Sounds (03020106), Upper Neuse (03020201), Middle Neuse (03020202), Contentnea (03020203), Lower Neuse (03020204), New (03030001), Haw (03030002), Deep (03030003), Upper Cape Fear (03030004), Lower Cape Fear (03030005), Black (03030006), Northeast Cape Fear (03030007), Upper Yadkin (03040101), South Yadkin (03040102), Lower Yadkin (03040103), Upper Pee Dee (03040104), Rocky, North Carolina, (03040105), Lower Pee Dee (03040201), Lynches (03040202), Lumber (03040203), Little Pee Dee (03040204), Black (03040205), Waccamaw (03040206), Carolina Coastal-Sampit (03040207), Upper Catawba (03050101), South Fork Catawba (03050102), Lower Catawba (03050103), Wateree (03050104), Upper Broad (03050105), Lower Broad (03050106), Tyger (03050107), Enoree (03050108), Saluda (03050109), Congaree (03050110), Lake Marion (03050111), Santee (03050112), Cooper (03050201), South Carolina Coastal (03050202), North Fork Edisto (03050203), South Fork Edisto (03050204), Edisto (03050205), Four Hole Swamp (03050206), Salkehatchie (03050207), Broad-St. Helena (03050208), Seneca (03060101), Tugaloo (03060102), Upper Savannah (03060103), Broad (03060104), Middle Savannah (03060106), Stevens (03060107)
04 Baptism-Brule (04010101), Beaver-Lester (04010102), St. Louis (04010201), Cloquet (04010202), Beartrap-Nemadji (04010301), Bad-Montreal (04010302), Black-Presque Isle (04020101), Ontonagon (04020102), Keweenaw Peninsula (04020103), Sturgeon (04020104), Dead-Kelsey (04020105), Betsy-Chocolay (04020201), Tahquamenon (04020202), Waiska (04020203), Lake Superior (04020300), Manitowoc-Sheboygan (04030101), Door-Kewaunee (04030102), Duck-Pensaukee (04030103), Oconto (04030104), Peshtigo (04030105), Brule (04030106), Michigamme (04030107), Menominee (04030108), Cedar-Ford (04030109), Escanaba (04030110), Tacoosh-Whitefish (04030111), Fishdam-Sturgeon (04030112), Upper Fox (04030201), Wolf (04030202), Lake Winnebago (04030203), Lower Fox (04030204), Little Calumet-Galien (04040001), Pike-Root (04040002), Milwaukee (04040003), St. Joseph (04050001), Black-Macatawa (04050002), Kalamazoo (04050003), Upper Grand (04050004), Maple (04050005), Lower Grand (04050006), Thornapple (04050007), Pere Marquette-White (04060101), Muskegon (04060102), Manistee (04060103), Betsie-Platte (04060104), Boardman-Charlevoix (04060105), Manistique (04060106), Brevoort-Millecoquins (04060107), Lake Michigan (04060200), St. Marys (04070001), Carp-Pine (04070002), Lone Lake-Ocqueoc (04070003), Cheboygan (04070004), Black (04070005), Thunder Bay (04070006), Au Sable (04070007), Au Gres-Rifle (04080101), Kawkawlin-Pine (04080102), Pigeon-Wiscoggin (04080103), Birch-Willow (04080104), Tittabawassee (04080201), Pine (04080202), Shiawassee (04080203), Flint (04080204), Cass (04080205), Saginaw (04080206), Lake Huron (04080300), St. Clair (04090001), Lake St. Clair (04090002), Clinton (04090003), Detroit (04090004), Huron (04090005), Ottawa-Stony (04100001), Raisin (04100002), St. Joseph (04100003), St. Marys (04100004), Upper Maumee (04100005), Tiffin (04100006), Auglaize (04100007), Blanchard (04100008)*, Lower Maumee (04100009), Cedar-Portage (04100010), Sandusky (04100011), Huron-Vermilion (04100012), Black-Rocky (04110001), Cuyahoga (04110002), Ashtabula-Chagrin (04110003), Grand (04110004), Chautauqua-Conneaut (04120101), Cattaraugus (04120102), Buffalo-Eighteenmile (04120103), Niagara (04120104), Lake Erie (04120200), Oak Orchard-Twelvemile (04130001), Upper Genesee (04130002), Lower Genesee (04130003), Irondequoit-Ninemile (04140101), Salmon-Sandy (04140102), Seneca (04140201), Oneida (04140202), Oswego (04140203), Black (04150101), Chaumont-Perch (04150102), Upper St. Lawrence (04150301), Oswegatchie (04150302), Indian (04150303), Grass (04150304), Raquette (04150305), St. Regis (04150306), English-Salmon (04150307)
05 Upper Allegheny (05010001), Conewango (05010002), Middle Allegheny-Tionesta (05010003), French (05010004), Clarion (05010005), Middle Allegheny-Redbank (05010006), Conemaugh (05010007), Lower Allegheny (05010009), Tygart Valley (05020001), West Fork (05020002), Upper Monongahela (05020003), Cheat (05020004), Lower Monongahela (05020005), Youghiogheny (05020006), Upper Ohio (05030101), Shenango (05030102), Mahoning (05030103)*, Beaver (05030104), Connoquenessing (05030105), Upper Ohio-Wheeling (05030106), Little Muskingum-Middle Island (05030201), Upper Ohio-Shade (05030202), Hocking (05030204), Tuscarawas (05040001), Mohican (05040002), Walhonding (05040003), Muskingum (05040004), Wills (05040005), Licking (05040006), Upper Kanawha (05050006), Lower Kanawha (05050008), Upper Scioto (05060001), Lower Scioto (05060002), Tug (05070201), Upper Great Miami (05080001), Lower Great Miami (05080002)*, Whitewater (05080003), Raccoon-Symmes (05090101), Little Scioto-Tygarts (05090103), Little Sandy (05090104), Ohio Brush-Whiteoak (05090201), Little Miami (05090202), Middle Ohio-Laughery (05090203), Licking (05100101), Upper Wabash (05120101), Salamonie (05120102), Mississinewa (05120103), Eel (05120104), Tippecanoe (05120106), Muscatatuck (05120207), Silver-Little Kentucky (05140101), Lower Ohio (05140206)
07 Mississippi Headwaters (07010101), Leech Lake (07010102), Prairie-Willow (07010103), Elk-Nokasippi (07010104), Pine (07010105), Crow Wing (07010106), Redeye (07010107), Long Prairie (07010108), Platte-Spunk (07010201), Sauk (07010202), Clearwater-Elk (07010203), Crow (07010204), South Fork Crow (07010205), Twin Cities (07010206), Rum (07010207), Upper Minnesota (07020001), Pomme De Terre (07020002), Lac Qui Parle (07020003), Hawk-Yellow Medicine (07020004), Chippewa (07020005), Redwood (07020006), Middle Minnesota (07020007), Cottonwood (07020008), Blue Earth (07020009), Watonwan (07020010), Le Sueur (07020011), Lower Minnesota (07020012), Upper St. Croix (07030001), Namekagon (07030002), Kettle (07030003), Snake (07030004), Lower St. Croix (07030005), Rush-Vermillion (07040001), Cannon (07040002), Buffalo-Whitewater (07040003), Zumbro (07040004), Trempealeau (07040005), La Crosse-Pine (07040006), Black (07040007), Root (07040008), Upper Chippewa (07050001), Flambeau (07050002), South Fork Flambeau (07050003), Jump (07050004), Lower Chippewa (07050005), Eau Claire (07050006), Red Cedar (07050007), Coon-Yellow (07060001), Grant-Little Maquoketa (07060003), Apple-Plum (07060005), Upper Wisconsin (07070001), Lake Dubay (07070002), Castle Rock (07070003), Baraboo (07070004), Lower Wisconsin (07070005), Kickapoo (07070006), Copperas-Duck (07080101), Upper Wapsipinicon (07080102)*, Lower Wapsipinicon (07080103)*, Flint-Henderson (07080104), South Skunk (07080105), Upper Cedar (07080201), Shell Rock (07080202), Winnebago (07080203), West Fork Cedar (07080204)*, Middle Cedar (07080205)*, Lower Cedar (07080206), Middle Iowa (07080208), Upper Rock (07090001), Crawfish (07090002), Pecatonica (07090003), Sugar (07090004), Lower Rock (07090005)*, Kishwaukee (07090006)*, Des Moines Headwaters (07100001), East Fork Des Moines (07100003), Boone (07100005), North Raccoon (07100006)*, South Raccoon (07100007)*, Lake Red Rock (07100008)*, Lower Des Moines (07100009)*, Kankakee (07120001), Iroquois (07120002), Chicago (07120003), Des Plaines (07120004), Upper Illinois (07120005), Upper Fox (07120006), Lower Fox (07120007)*, Lower Illinois-Senachwine Lake (07130001)*, Lower Illinois-Lake Chautauqua (07130003)*, Lower Illinois (07130011)*, Meramec (07140102)*, Lower Kaskaskia (07140204)*
09 Bois De Sioux (09020101), Mustinka (09020102), Otter Tail (09020103), Upper Red (09020104), Western Wild Rice (09020105), Buffalo (09020106), Elm-Marsh (09020107), Eastern Wild Rice (09020108), Goose (09020109), Lower Sheyenne (09020204), Sandhill-Wilson (09020301), Red Lakes (09020302), Red Lake (09020303), Thief (09020304), Clearwater (09020305), Grand Marais-Red (09020306), Turtle (09020307), Forest (09020308), Snake (09020309), Park (09020310), Lower Red (09020311), Two Rivers (09020312), Roseau (09020314), Rainy Headwaters (09030001), Vermilion (09030002), Rainy Lake (09030003), Upper Rainy (09030004), Little Fork (09030005), Big Fork (09030006), Rapid (09030007), Lower Rainy (09030008), Lake of the Woods (09030009)
10 Upper Big Sioux (10170202), Lower Big Sioux (10170203), Rock (10170204), Blackbird-Soldier (10230001)*, Little Sioux (10230003), Boyer (10230007), Nodaway (10240010)*, Thompson (10280102)*, Lower Grand (10280103)
+ Natural heritage record(s) exist for this watershed
* Extirpated/possibly extirpated
U.S. Distribution by Watershed (based on multiple information sources) Help
Ecology & Life History
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Reproduction Comments: Spawns in spring and summer; eggs hatch in about 3-5 days; male guards eggs; sexually mature in 2nd or 3rd year; may nest in colonies (Moyle 1976).
Habitat Type: Freshwater
Non-Migrant: Y
Locally Migrant: N
Long Distance Migrant: N
Riverine Habitat(s): CREEK, Low gradient, MEDIUM RIVER
Lacustrine Habitat(s): Shallow water
Palustrine Habitat(s): HERBACEOUS WETLAND
Special Habitat Factors: Benthic
Habitat Comments: Lakes, reservoirs, ponds, sloughs, and sluggish streams; prefers quiet, clear water with aquatic vegetation and some organic debris. May occur in large numbers in shallow sheltered situations. Eggs are laid in a nest constructed by the male in shallow water (less than 1 m deep) in bottoms of sand, gravel, or woody debris.
Adult Food Habits: Invertivore
Immature Food Habits: Invertivore
Food Comments: Eats snails, aquatic insects, and other invertebrates obtained from bottom or from plant surfaces; pharyngeal jaws are specialized for crushing hard-shelled prey; larvae eat zooplankton (Moyle 1976, Becker 1983).
Length: 40 centimeters
Economic Attributes Not yet assessed
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Management Summary Not yet assessed
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Population/Occurrence Delineation
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Group Name: Sunfishes (Centrarchids)

Use Class: Not applicable
Minimum Criteria for an Occurrence: Occurrences are based on evidence of historical presence, or current and likely recurring presence, at a given location. Such evidence minimally includes collection or reliable observation and documentation of one or more individuals (including eggs and larvae) in appropriate habitat.
Separation Barriers: Dam lacking a suitable fishway; high waterfall; upland habitat.
Separation Distance for Unsuitable Habitat: 10 km
Separation Distance for Suitable Habitat: 10 km
Separation Justification: Separation distance is arbitrary. Although members of this group vary in size and probably in typical movement distances, it is likely that even the smallest centrarchids occasionally disperse as far as do large centrarchids. Hence a single separation distance is used for all members of the family. Because of the difficulty in defining suitable versus unsuitable habitat, especially with respect to dispersal, and to simplify the delineation of occurrences, a single separation distance is used regardless of habitat quality.
Date: 25Jun2001
Author: Hammerson, G.
Notes: Note that some species some species may at time be hard to detect. For example, nowhere is the Carolina pygmy sunfish known to be abundant. In addition, it is essentially an annual species, with adults dying soon after spawning, at an age of 12-15 months. In addition, young are so small that, for a several months, documentation of the species' presence at a particular locality might be almost impossible, at least without preserving specimens. Therefore, negative data at a known locality should be carefully interpreted (P. Shute).
Population/Occurrence Viability
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U.S. Invasive Species Impact Rank (I-Rank) Not yet assessed
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Authors/Contributors
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NatureServe Conservation Status Factors Edition Date: 07Aug2012
Element Ecology & Life History Edition Date: 07Oct1993
Element Ecology & Life History Author(s): Hammerson, G.

Zoological data developed by NatureServe and its network of natural heritage programs (see Local Programs) and other contributors and cooperators (see Sources).

References
Help
  • Aquin, P. 1999. Évaluation de la situation des groupes taxonomiques des poissons du Québec. Ministère de l'Environnement et de la Faune. 9 pages.

  • Burr, B. M., and P. W. Smith. 1976. Status of the largescalestoneroller, Campostoma oligolepis. Copeia 1976(3):521-531.

  • Lee, D. S., C. R. Gilbert, C. H. Hocutt, R. E. Jenkins, D. E. McAllister, and J. R. Stauffer, Jr. 1980. Atlas of North American freshwater fishes. North Carolina State Museum of Natural History, Raleigh, North Carolina. i-x + 854 pp.

  • Legendre, V. et J.F. Bergeron. 1977. Liste des poissons d' eau douce du Québec. MLCP, Service Aménage. Expl. Faune. Rap. dact. 6

  • Manooch, C. S., III. 1984. Fisherman's guide. Fishes of the southeastern United States. North Carolina State Museum of Natural History, Raleigh. 362 pp.

  • Moyle, P. B. 1976a. Inland fishes of California. University of California Press, Berkeley, California. 405 pp.

  • Moyle, P. B. 2002. Inland fishes of California. Revised and expanded. University of California Press, Berkeley. xv + 502 pp.

  • Nelson, J. S., E. J. Crossman, H. Espinosa-Perez, L. T. Findley, C. R. Gilbert, R. N. Lea, and J. D. Williams. 2004. Common and scientific names of fishes from the United States, Canada, and Mexico. American Fisheries Society, Special Publication 29, Bethesda, Maryland. 386 pp.

  • Page, L. M., H. Espinosa-Pérez, L. T. Findley, C. R. Gilbert, R. N. Lea, N. E. Mandrak, R. L. Mayden, and J. S. Nelson. 2013. Common and scientific names of fishes from the United States, Canada, and Mexico. Seventh edition. American Fisheries Society, Special Publication 34, Bethesda, Maryland.

  • Page, L. M., and B. M. Burr. 2011. Peterson field guide to freshwater fishes of North America north of Mexico. Second edition. Houghton Mifflin Harcourt, Boston. xix + 663 pp.

  • Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea, and W.B. Scott. 1991. Common and scientific names of fishes from the United States and Canada. American Fisheries Society, Special Publication 20. 183 pp.

References for Watershed Distribution Map
  • Becker, G. C. 1983. Fishes of Wisconsin. University of Wisconsin Press, Madison. 1,052 pp.

  • Burr, B. M., and M. L. Warren, Jr. 1986a. Distributional atlas of Kentucky fishes. Kentucky Nature Preserves Commission, Scientific and Technical Series No. 4, Frankfort, Kentucky. 398 pp.

  • Cooper, E. L. 1983. Fishes of Pennsylvania and the northeastern United States. Pennsylvania State University Press, University Park. 243 pp.

  • Fago, D. 2000. Relative abundance and distribution of fishes in Wisconsin. Fish Distribution Database to year 2000. Wisconsin Department of Natural Resources.

  • Harlan, J. R., E. B. Speaker, and J. Mayhew. 1987. Iowa fish and fishing. Iowa Conservation Commission, Des Moines, Iowa. 323 pp.

  • Jenkins, R. E., and N. M. Burkhead. 1994. Freshwater fishes of Virginia. American Fisheries Society, Bethesda, Maryland. xxiii + 1079 pp.

  • Marcy, B. C., Jr., D. E. Fletcher, F. D. Martin, M. H. Paller, and M.J.M. Reichert. 2005. Fishes of the middle Savannah River basin. University of Georgia Press, Athens. xiv + 460 pp.

  • Menhinick, E. F. 1991. The freshwater fishes of North Carolina. North Carolina Wildlife Resources Commission. 227 pp.

  • Page, L. M., and B. M. Burr. 1991. A field guide to freshwater fishes: North America north of Mexico. Houghton Mifflin Company, Boston, Massachusetts. 432 pp.

  • Pflieger, W. L. 1975. The fishes of Missouri. Missouri Department of Conservation. Columbia, Missouri. viii + 343 pp.

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  • Smith, P. W. 1979. The fishes of Illinois. University of Illinois Press, Urbana. 314 pp.

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  • Whitworth, W. R., P. L. Berrien, and W. T. Keller. 1976. Freshwater fishes of Connecticut. Bulletin of the Connecticut Geological and Natural History Survey 101. vi + 134 pp.

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