Cambarellus diminutus - Hobbs, 1945
Least Crayfish
Taxonomic Status: Accepted
Related ITIS Name(s): Cambarellus diminutus Hobbs, 1945 (TSN 97620)
Unique Identifier: ELEMENT_GLOBAL.2.119084
Element Code: ICMAL09020
Informal Taxonomy: Animals, Invertebrates - Crustaceans - Crayfishes
 
Kingdom Phylum Class Order Family Genus
Animalia Crustacea Malacostraca Decapoda Cambaridae Cambarellus
Genus Size: C - Small genus (6-20 species)
Check this box to expand all report sections:
Concept Reference
Help
Concept Reference: Hobbs, H.H., Jr. 1989. An illustrated checklist of the American crayfishes (Decapoda: Astacidae, Cambaridae, and Parastacidae). Smithsonian Contributions to Zoology 480:1-236.
Concept Reference Code: B89HOB01EHUS
Name Used in Concept Reference: Cambarellus diminutus
Conservation Status
Help

NatureServe Status

Global Status: G3
Global Status Last Reviewed: 17May2010
Global Status Last Changed: 19Feb1996
Rounded Global Status: G3 - Vulnerable
Reasons: It has a small extent of occurrence (<8000 sq. km) and has a continuing decline in habitat extent and quality due to dockland usage, agricultural intensification and urbanization. Even though this species can be considered a generalist, its preference for temporary habitats consisting of thick vegetation, leaves it sensitive to any habitat alterations. Further, with the continued pollution associated with docklands and agriculture, it is not known when this will start to severely impact populations. Due to the intensification of agriculture, the likelihood of oxbow side ponds remaining long enough to be utilized by this species is uncertain.
Nation: United States
National Status: N3 (19Feb1996)

U.S. & Canada State/Province Status
Due to latency between updates made in state, provincial or other NatureServe Network databases and when they appear on NatureServe Explorer, for state or provincial information you may wish to contact the data steward in your jurisdiction to obtain the most current data. Please refer to our Distribution Data Sources to find contact information for your jurisdiction.
United States Alabama (S2), Mississippi (S2)

Other Statuses

IUCN Red List Category: DD - Data deficient
American Fisheries Society Status: Threatened (01Aug2007)

NatureServe Global Conservation Status Factors

Range Extent: 5000-20,000 square km (about 2000-8000 square miles)
Range Extent Comments: Apparently confined to Mobile County, Alabama, and George and Jackson counties, Mississippi (Hobbs, 1989).

Number of Occurrences: 6 - 80
Number of Occurrences Comments: The total number of occurences will eventually prove to be very close to the B-C line of demarcation; presently ca. 20 sites. In the original description, Hobbs (1945) cites Mobile Co. (stream south of Irvington; and a ditch west of Grand Bay), Alabama, and Jackson Co. (burrow west of Grand Bay, Alabama), Mississippi. In Alabama, known from eight records, all from Mobile and Washington Cos. in the Mobile and Pascagoula basins (Mirarchi et al., 2004; appendix 1-2 pub. separately; Schuster and Taylor, 2004; Schuster et al., 2008). Peterson et al. (1996) found the species in 3 of 7 surveyed drainages (9 sites in upper Bluff Creek, Old Ft. Bayou, pond in Tchoutacabouffa River) along the Mississippi Gulf coast; with all occurring in the more eastern drainages.

Population Size: 10,000 to >1,000,000 individuals
Population Size Comments: Although the numbers are high (maybe even D status), because of the very small size, the biomass is low

Overall Threat Impact Comments: It is unknown whether it is impacted by any major threat process. However, Mobile county is the third most populated city in the southern USA, consisting of a large dockland areas which is intensively used. This will be creating a substantial water pollution in this area of Mobile county, whilst also polluting surrounding streams and ponds. Further, C. diminutus' use of oxbow side ponds is useful for protection against predation, but these are only temporary habitats and therefore there must be other similar habitats avaliable for this species to utilize once the oxbow side ponds have silted up (Peterson et al., 1996). However, this species is somewhat tolerant of differing water and habitat qualities to exist in these habitats (Hobbs, 1945). But, with the continued expansion of the Mobile docklands usage and intensive agriculture, it is unknown how long it will be able to use these habitats (S. Adams, pers. comm., 2009).

Short-term Trend: Decline of 10-30%
Short-term Trend Comments: It has a small extent of occurrence (<8000 sq. km) and has a continuing decline in habitat extent and quality due to dockland usage, agricultural intensification and urbanization.

Intrinsic Vulnerability Comments: Most species of the subgenus are resilient and rebound well

Other NatureServe Conservation Status Information

Inventory Needs: population size estimates

Distribution
Help
Global Range: (5000-20,000 square km (about 2000-8000 square miles)) Apparently confined to Mobile County, Alabama, and George and Jackson counties, Mississippi (Hobbs, 1989).

U.S. States and Canadian Provinces

Due to latency between updates made in state, provincial or other NatureServe Network databases and when they appear on NatureServe Explorer, for state or provincial information you may wish to contact the data steward in your jurisdiction to obtain the most current data. Please refer to our Distribution Data Sources to find contact information for your jurisdiction.
Color legend for Distribution Map
Endemism: endemic to a single nation

U.S. & Canada State/Province Distribution
United States AL, MS

Range Map
No map available.


U.S. Distribution by County Help
State County Name (FIPS Code)
AL Mobile (01097), Washington (01129)
MS George (28039)*, Jackson (28059)
* Extirpated/possibly extirpated
U.S. Distribution by Watershed Help
Watershed Region Help Watershed Name (Watershed Code)
03 Lower Tambigbee (03160203)+, Pascagoula (03170006)+*, Escatawpa (03170008)+, Mississippi Coastal (03170009)+
+ Natural heritage record(s) exist for this watershed
* Extirpated/possibly extirpated
Ecology & Life History
Help
Basic Description: a crayfish
General Description: Hooks on ischia of male third and fourth pereiopods; male pleopod with three terminal elements, all curved at about 45 degrees to main axis of pleopod, caudal process broadly triangular; cervical spines strong (Fitzpatrick, 1983). [LENGTH: to 12 TCL, to 30 TL] [WIDTH: to 6]
Diagnostic Characteristics: Hooks on 3rd & 4th pereiopods, terminal elements short, coarser than typical in subgenus, with caudal process broadly triangular.
Reproduction Comments: No empirical data, but broad seasonal occurrence of ovigerous females implies year round breeding.
Ecology Comments: See GSPECFACTOR & GHABCOM.
Habitat Type: Freshwater
Non-Migrant: N
Locally Migrant: N
Long Distance Migrant: N
Mobility and Migration Comments: No data; home range probably does not exceed 20 m dia.
Riverine Habitat(s): CREEK, Low gradient, Pool
Lacustrine Habitat(s): Shallow water
Special Habitat Factors: Benthic, Burrowing in or using soil
Habitat Comments: Found among vegetation in small to moderate blackwater streams, usually in pine woods. Also invades ditches that communicate with sluggish streams; tolerant of warm water, but prefers shaded areas (Hobbs, 1989). Seems to prefer very sluggish flow with submerged, at least in part, vegetation. Will burrow during dry conditions.
Adult Food Habits: Detritivore
Immature Food Habits: Detritivore
Food Comments: No data; probably opportunistic, mostly detritus.
Adult Phenology: Circadian
Immature Phenology: Circadian
Economic Attributes
Help
Economic Comments: No known economic value to humans.
Management Summary
Help
Biological Research Needs: fecundity
Population/Occurrence Delineation
Help
Group Name: Crayfishes

Use Class: Not applicable
Minimum Criteria for an Occurrence: Occurrences are based on some evidence of historical or current presence of single or multiple specimens, including live specimens or recently dead shells (i.e., soft tissue still attached without signs of external weathering or staining), at a given location with potentially recurring existence. Evidence is derived from reliable published observation or collection data; unpublished, though documented (i.e. government or agency reports, web sites, etc.) observation or collection data; or museum specimen information.
Separation Barriers: Separation barriers are based on hydrological discontinuity. Additional physical barriers, particularly for secondary and tertiary burrowers, include presence of upland habitat between water connections of a distance greater than 30 m. Migration of primary burrowers is generally not hindered by presence of upland habitat unless conditions are very xeric (dry and desert-like) (Smith, 2001).
Separation Distance for Unsuitable Habitat: 2 km
Separation Distance for Suitable Habitat: 2 km
Alternate Separation Procedure: Freshwater cave (troglobitic) species may occur from near entrances to very deep in cave systems. For cave species, each cave where an observation or collection was recorded (see Minimum EO Criteria, above) constitutes an element occurrence regardless of separation distance unless caves are part of a single hydrological system (see below). Occurrences are additionally separated by underground physical barriers to movement. Multiple caves within a single hydrological cave system are considered to be a single element occurrence when they are less than one km apart. Multiple caves within a single hydrological cave system are considered separate element occurrences when hydrological connections have not been determined or when separated by a distance of at least one km.
Separation Justification: Habitat for these creatures is primarily separated according to each species' burrowing ability. All crayfish are able to burrow to some extent and this ability will help determine the range of habitats in which a species can be found. Burrowing in the Astacidae is limited to streambed and bank excavation (Hobbs, 1988). The Cambaridae, as a whole are much more adept at burrowing than the Astacidae. As a result, they possess a greater habitat range than the Astacidae including dry water bodies (Hogger, 1988).

The burrowers can be classified into three categories: primary burrowers, secondary burrowers, and tertiary burrowers. Primary burrowers tend to remain in their burrows continuously and live in areas without permanent water except during breeding when they must migrate to a nearby water source (Hogger, 1988). The prairies of eastern and central Mississippi and western Alabama are an example of primary burrower habitat (Hogger, 1988). Secondary burrowers remain in burrows during dry periods but emerge when habitats are inundated seasonally. Such habitat includes lentic systems flooded periodically but dry in summer (Huner and Romaire, 1979) and permanent and temporary ponds and swamps in the southern United States. Tertiary burrowers do not burrow except during infrequent drought conditions and/or during breeding season. Both flowing and standing water can be tertiary burrower habitat.

Because primary burrowers, and to a lesser extent secondary burrowers, can occupy xeric habitats, separation barriers for such species do not include presence of upland habitat except in extremely dry conditions. Survival during dry periods, particularly for secondary burrowers, is dependent upon construction of a burrow regardless of season. Several different types have been described (Smith, 2001) depending on species, soil, and depth of water table.

Published information about movement in relation to migration distance is lacking but Cooper (1998, personal communication) and Fitzpatrick (1998, personal communication) both recommend a separation distance of one km between element occurrences. Dispersal patterns are best known for invasive species which likely have the greatest dispersal capability, therefore, separation distances have been determined for all crayfish based on these studies. Guan and Wiles (1997) provided evidence from the River Great Ouse in the United Kingdom that the range of movement for the majority of the invasive Pacifastacus leniusculus was within 190 m. Bubb et al. (2004) also studied P. leniusculus in England using radio-tagging and found median maximal upstream and downstream movement distances were 13.5 m (range 0-283 m) and 15 m (range 0-417 m), respectively. Barbaresi et al. (2004) found that ranging speed in the invasive crayfish Procambarus clarkii (Girard) to be slow (0.3 to 76.5 m/day) with the widest ranging individual traveling 304 m. Lewis and Horton (1996) found that 21% of tagged Pacifastacus leniusculus in an Oregon harvest pond moved >1000 m in one year while the majority moved <500 m. As such minimum separation distance (unsuitable and suitable) has been set at the NatureServe standard minimum of two km.

Exposed pools and streams in caves represent "karst windows" into more extensive underground streams. No information on the distance cave crayfish can disperse in underground streams is yet available.

Date: 18Oct2004
Author: Cordeiro, J.
Notes: Primary burrowers include the following taxa: Cambarus (Cambarus) carolinus, C. (C.) diogenes diogenes, C. (Depressicambarus) catagius, C. (D.) cymatilis, C. (D.) deweesae, C. (D.) harti, C. (D.) reflexus, C. (D.) pyronotus, C. (D.) striatus, C. (D.) strigosus, C. (D.) truncatus, C. (Glareocola), C. (Jugicambarus) batchi, C. (J.) carolinus, C. (J.) causeyi, C. (J.) dubius, C. (J.) gentryi, C. (J.) monongalensis, C. (J.) nodosus, C. (Lacunicambarus), C. (Tubericambarus), Distocambarus, Fallicambarus, Procambarus (Acucauda), P. (Distocambarus), P. (Girardiella) barbiger, P. (G.) cometes, P. (G.) connus, P. (G.) curdi, P. (G.) gracilis, P. (G.) hagenianus hagenianus, P. (G.) hagenianus vesticeps, P. (G.) liberorum, P. (G.) pogum, P. (Hagenides) [except P. pygmaeus]
Secondary burrowers include the following taxa: Cambarus (Cambarus) ortmanni, C. (Depressicambarus) latimanus, C. (D.) reduncus, Hobbseus, Procambarus (Cambarus) clarkii, P. (Girardiella) kensleyi, P. (G.) reimeri, P. (G.) simulans, P. (G.) steigmani, P. (G.) tulanei, P. (Hagenides) pygmaeus, P. (Leconticambarus) [excepting P. alleni and P. milleri], P. (Ortmannicus) [excepting the cave dwelling species], P. (Tenuicambarus)
Tertiary burrowers include the following taxa: Barbicambarus, Bouchardina, Cambarus (Cambarus) angularis, C. (C.) bartonii carinirostris, C. (C.) bartonii cavatus, C. (C.) howardi, C. (C.) sciotensis, C. (Depressicambarus) englishi, C. (D.) graysoni, C. (D.) halli, C. (D.) obstipus, C. (D.) sphenoides, C. (Erebicambarus) ornatus, C. (E.) rusticiformis, C. (Exilicambarus) cracens, C. (Hiaticambarus), C. (Jugicambarus) asperimanus, C. (J.) bouchardi, C. (J.) crinipes, C. (J.) distans, C. (J.) friaufi, C. (J.) obeyensis, C. (J.) parvoculus, C. (J.) unestami, C. (Puncticambarus) [excepting the cave dwelling species], C. (Veticambarus), Cambarellus, Faxonella, Orconectes [excepting the cave dwelling species], Pacifastacus, Procambarus (Capillicambarus), P. (Girardiella) ceruleus, P.

Population/Occurrence Viability
Help
U.S. Invasive Species Impact Rank (I-Rank) Not yet assessed
Help
Authors/Contributors
Help
NatureServe Conservation Status Factors Edition Date: 17May2010
NatureServe Conservation Status Factors Author: Cordeiro, J. (2010); Fitzpatrick, J.F., Jr. (1989)
Element Ecology & Life History Edition Date: 12Sep2008
Element Ecology & Life History Author(s): Cordeiro, J. (2008); FITZPATRICK, J.F. (1991)

Zoological data developed by NatureServe and its network of natural heritage programs (see Local Programs) and other contributors and cooperators (see Sources).

References
Help
  • Alabama Department of Conservation and Natural Resources, Division of Wildlife and Freshwater Fisheries. 2005. Conserving Alabama's wildlife: a comprehensive strategy. Alabama Department of Conservation and Natural Resources, Division of Wildlife and Freshwater Fisheries. Montgomery, Alabama. 303 pages. [Available online at http://www.dcnr.state.al.us/research-mgmt/cwcs/outline.cfm ]

  • Fitzpatrick, J.F., Jr. 1983. A revision of the dwarf crawfishes (Cambaridae, Cambarellinae). Journal of Crustacean Biology 3(2):266-277.

  • Hobbs, H.H., Jr. 1945. Two new species of crayfishes of the genus Cambarellus from the Gulf coastal states, with a key to the species of the genus (Decapoda, Astacidae). American Midland Naturalist 34(2):466-474.

  • Hobbs, H.H., Jr. 1989. An illustrated checklist of the American crayfishes (Decapoda: Astacidae, Cambaridae, and Parastacidae). Smithsonian Contributions to Zoology 480:1-236.

  • McLaughlin, P.A., D.K. Camp, M.V. Angel, E.L. Bousfield, P. Brunel, R.C. Brusca, D. Cadien, A.C. Cohen, K. Conlan, L.G. Eldredge, D.L. Felder, J.W. Goy, T. Haney, B. Hann, R.W. Heard, E.A. Hendrycks, H.H. Hobbs III, J.R. Holsinger, B. Kensley, D.R. Laubitz, S.E. LeCroy, R. Lemaitre, R.F. Maddocks, J.W. Martin, P. Mikkelsen, E. Nelson, W.A. Newman, R.M. Overstreet, W.J. Poly, W.W. Price, J.W. Reid, A. Robertson, D.C. Rogers, A. Ross, M. Schotte, F. Schram, C. Shih, L. Watling, G.D.F. Wilson, and D.D. Turgeon. 2005. Common and scientific names of aquatic invertebrates from the United States and Canada: crustaceans. American Fisheries Society Special Publication 31: 545 pages.

  • McLaughlin, P.A., D.K. Camp, M.V. Angel, E.L. Bousfield, P. Brunel, R.C. Brusca, D. Cadien, A.C. Cohen, K. Conlan, L.G. Eldredge, D.L. Felder, J.W. Goy, T. Haney, B. Hann, R.W. Heard, E.A. Hendrycks, H.H. Hobbs III, J.R. Holsinger, B. Kensley, D.R. Laubitz, S.E. LeCroy, R. Lemaitre, R.F. Maddocks, J.W. Martin, P. Mikkelsen, E. Nelson, W.A. Newman, R.M. Overstreet, W.J. Poly, W.W. Price, J.W. Reid, A. Robertson, D.C. Rogers, A. Ross, M. Schotte, F. Schram, C. Shih, L. Watling, G.D.F. Wilson, and D.D. Turgeon. 2005. Common and scientific names of aquatic invertebrates from the United States and Canada: Crustaceans. American Fisheries Society Special Publication 31: 545 pp.

  • Mirarchi, R.E., M.A. Bailey, J.T. Garner, T.M. Haggerty, T.L. Best, M.F. Mettee, and P. O'Neil. 2004d. Alabama Wildlife. Volume Four: Conservation and Management Recommendations for Imperiled Wildlife. University of Alabama Press, Tuscaloosa, Alabama. 221 pp.

  • Peterson, M. S., J. F. Fitzpatrick, Jr., and S. J. VanderKooy. 1996. Distribution and habitat use by dwarf crayfishes (Decapoda: Cambaridae: Cambarellus). Wetlands 16:594-598.

  • Schuster, G. A. and C.A. Taylor. 2004. Report on the crayfishes of Alabama: literature and museum database review, species list with abbreviated annotations and proposed conservation statuses. Illinois Natural History Survey Technical Report, 2004(12). Report submitted to the Alabama Department of Conservation and Natural Resources, Wildlife and Freshwater Fisheries Division, Montgomery, Alabama. 47 pages.

  • Schuster, G. A. and C.A. Taylor. 2004. Report on the crayfishes of Alabama: literature and museum database review, species list with abbreviated annotations and proposed conservation statuses. Illinois Natural History Survey Technical Report, 2004(12): 47 pp.

  • Schuster, G.A., C.A. Taylor, and J. Johansen. 2008. An annotated checklist and preliminary designation of drainage distributions of the crayfishes of Alabama. Southeastern Naturalist, 7(3): 493-504.

  • Smith, J. Brett, Guenter A. Schuster, Christopher A. Taylor, E. Anne Wynn, and Stuart W. McGregor. 2011. A preliminary report on the distribution and conservation status of the Alabama crayfish fauna. Geologic Survey of Alabama Open-file Report 1102. Tuscaloosa Alabama. 180 pages.

  • Taylor, C.A., G.A. Schuster, J.E. Cooper, R.J. DiStefano, A.G. Eversole, P. Hamr, H.H. Hobbs III, H.W. Robison, C.E. Skelton, and R.F. Thoma. 2007. A reassessment of the conservation status of crayfishes of the United States and Canada after 10+ years of increased awareness. Fisheries 32(8):371-389.

  • Taylor, Christopher A. and Guenter A. Schuster. 2007. Final report: compilation of Alabama crayfish museum holdings and construction of a geo-referenced database. Illinois Natural Histroy Survey, Center for Biodiversity and Ecological Entomology Technical Report 2007(26). 14 pages.

Use Guidelines & Citation

Use Guidelines and Citation

The Small Print: Trademark, Copyright, Citation Guidelines, Restrictions on Use, and Information Disclaimer.

Note: All species and ecological community data presented in NatureServe Explorer at http://explorer.natureserve.org were updated to be current with NatureServe's central databases as of March 2018.
Note: This report was printed on

Trademark Notice: "NatureServe", NatureServe Explorer, The NatureServe logo, and all other names of NatureServe programs referenced herein are trademarks of NatureServe. Any other product or company names mentioned herein are the trademarks of their respective owners.

Copyright Notice: Copyright © 2018 NatureServe, 4600 N. Fairfax Dr., 7th Floor, Arlington Virginia 22203, U.S.A. All Rights Reserved. Each document delivered from this server or web site may contain other proprietary notices and copyright information relating to that document. The following citation should be used in any published materials which reference the web site.

Citation for data on website including State Distribution, Watershed, and Reptile Range maps:
NatureServe. 2018. NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. NatureServe, Arlington, Virginia. Available http://explorer.natureserve.org. (Accessed:

Citation for Bird Range Maps of North America:
Ridgely, R.S., T.F. Allnutt, T. Brooks, D.K. McNicol, D.W. Mehlman, B.E. Young, and J.R. Zook. 2003. Digital Distribution Maps of the Birds of the Western Hemisphere, version 1.0. NatureServe, Arlington, Virginia, USA.

Acknowledgement Statement for Bird Range Maps of North America:
"Data provided by NatureServe in collaboration with Robert Ridgely, James Zook, The Nature Conservancy - Migratory Bird Program, Conservation International - CABS, World Wildlife Fund - US, and Environment Canada - WILDSPACE."

Citation for Mammal Range Maps of North America:
Patterson, B.D., G. Ceballos, W. Sechrest, M.F. Tognelli, T. Brooks, L. Luna, P. Ortega, I. Salazar, and B.E. Young. 2003. Digital Distribution Maps of the Mammals of the Western Hemisphere, version 1.0. NatureServe, Arlington, Virginia, USA.

Acknowledgement Statement for Mammal Range Maps of North America:
"Data provided by NatureServe in collaboration with Bruce Patterson, Wes Sechrest, Marcelo Tognelli, Gerardo Ceballos, The Nature Conservancy-Migratory Bird Program, Conservation International-CABS, World Wildlife Fund-US, and Environment Canada-WILDSPACE."

Citation for Amphibian Range Maps of the Western Hemisphere:
IUCN, Conservation International, and NatureServe. 2004. Global Amphibian Assessment. IUCN, Conservation International, and NatureServe, Washington, DC and Arlington, Virginia, USA.

Acknowledgement Statement for Amphibian Range Maps of the Western Hemisphere:
"Data developed as part of the Global Amphibian Assessment and provided by IUCN-World Conservation Union, Conservation International and NatureServe."

NOTE: Full metadata for the Bird Range Maps of North America is available at:
http://www.natureserve.org/library/birdDistributionmapsmetadatav1.pdf.

Full metadata for the Mammal Range Maps of North America is available at:
http://www.natureserve.org/library/mammalsDistributionmetadatav1.pdf.

Restrictions on Use: Permission to use, copy and distribute documents delivered from this server is hereby granted under the following conditions:
  1. The above copyright notice must appear in all copies;
  2. Any use of the documents available from this server must be for informational purposes only and in no instance for commercial purposes;
  3. Some data may be downloaded to files and altered in format for analytical purposes, however the data should still be referenced using the citation above;
  4. No graphics available from this server can be used, copied or distributed separate from the accompanying text. Any rights not expressly granted herein are reserved by NatureServe. Nothing contained herein shall be construed as conferring by implication, estoppel, or otherwise any license or right under any trademark of NatureServe. No trademark owned by NatureServe may be used in advertising or promotion pertaining to the distribution of documents delivered from this server without specific advance permission from NatureServe. Except as expressly provided above, nothing contained herein shall be construed as conferring any license or right under any NatureServe copyright.
Information Warranty Disclaimer: All documents and related graphics provided by this server and any other documents which are referenced by or linked to this server are provided "as is" without warranty as to the currentness, completeness, or accuracy of any specific data. NatureServe hereby disclaims all warranties and conditions with regard to any documents provided by this server or any other documents which are referenced by or linked to this server, including but not limited to all implied warranties and conditions of merchantibility, fitness for a particular purpose, and non-infringement. NatureServe makes no representations about the suitability of the information delivered from this server or any other documents that are referenced to or linked to this server. In no event shall NatureServe be liable for any special, indirect, incidental, consequential damages, or for damages of any kind arising out of or in connection with the use or performance of information contained in any documents provided by this server or in any other documents which are referenced by or linked to this server, under any theory of liability used. NatureServe may update or make changes to the documents provided by this server at any time without notice; however, NatureServe makes no commitment to update the information contained herein. Since the data in the central databases are continually being updated, it is advisable to refresh data retrieved at least once a year after its receipt. The data provided is for planning, assessment, and informational purposes. Site specific projects or activities should be reviewed for potential environmental impacts with appropriate regulatory agencies. If ground-disturbing activities are proposed on a site, the appropriate state natural heritage program(s) or conservation data center can be contacted for a site-specific review of the project area (see Visit Local Programs).

Feedback Request: NatureServe encourages users to let us know of any errors or significant omissions that you find in the data through (see Contact Us). Your comments will be very valuable in improving the overall quality of our databases for the benefit of all users.