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Amphianthus pusillus - Isoetes melanospora - Isoetes tegetiformans Granite Pool Vegetation
Translated Name: Poolsprite - Blackspore Quillwort - Merlin's-grass Granite Pool Vegetation
Common Name: Piedmont Granitic Flatrock Glade, Seasonal Pool
Unique Identifier: CEGL004342
Classification Approach: International Vegetation Classification (IVC)
Summary: This community occupies the deepest pools with the longest hydroperiods found on granitic flatrocks of Georgia, South Carolina, and Alabama. These pools have intact rims, shallow soils, and are of sufficient depth to retain water for several weeks after a rain. The vegetation of these pools is limited to the rare aquatic endemics Amphianthus pusillus, Isoetes melanospora, and Isoetes tegetiformans.



Classification

Classification Confidence: Low - Poorly Documented
Classification Comments: Coffey (1964) cites Isoetes melanopoda as occurring "in crevices and depressions where water collects from the winter rains", with and Amphianthus pusillus growing "in abundance around the edges of the Isoetes patches." She does not name or identify either particular zones or specific associations in her work on flat granitic outcrops.

Vegetation Hierarchy
Class 2 - Shrub & Herb Vegetation
Subclass 2.C - Shrub & Herb Wetland
Formation 2.C.4 - Temperate to Polar Freshwater Marsh, Wet Meadow & Shrubland
Division 2.C.4.Nd - Eastern North American Temperate & Boreal Freshwater Marsh, Wet Meadow & Shrubland
Macrogroup Eastern North American Marsh, Wet Meadow & Shrubland
Group Southeastern Granite Outcrop Pool
Alliance Granitic Outcrop Pool

This is the revised vegetation hierarchy. For more information see Classification Sources and usnvc.org.

Similar Associations
Unique Identifier Name
CEGL004298 Packera tomentosa - Croton willdenowii - Schizachyrium scoparium - (Selaginella rupestris) Flatrock Vegetation
CEGL004344 Diamorpha smallii - Minuartia glabra - Minuartia uniflora - Cyperus granitophilus Flatrock Vegetation
CEGL007083 Isoetes lithophila - Isoetes melanopoda - Crassula aquatica - Callitriche peploides Granite Pool Vegetation



Related Concepts from Other Classifications

Related Subnational Community Units
These data are subject to substantial ongoing revision and may be out of date for some states.
In the U.S., contact the state Heritage Program for the most complete and up-to-date information at: http://www.natureserve.org/natureserve-network.
Information from programs in other jurisdictions will be posted when they are made available.
Subnation Concept Name Relationship to Standard Confidence Reference
Alabama Amphianthus pusillus - Isoetes melanospora - Isoetes tegetiformans Herbaceous Vegetation Equivalent Certain Schotz pers. comm.
South Carolina Granitic flatrock Broader   Nelson 1986


Other Related Concepts
Related Concept Name: Pool Type A
Relationship: = - Equivalent
Reference: Baker, W. B. 1956. Some interesting plants of the granite outcrops of Georgia. Georgia Mineral Newsletter 9(1):10-19.
Related Concept Name: shallow pool communities
Relationship: ? - Undetermined
Reference: Quarterman, E., M. P. Burbanck, and D. J. Shure. 1993. Rock outcrop communities: Limestone, sandstone, and granite. Pages 35-86 in: W. H. Martin, S. G. Boyce, and A. C. Echternacht, editors. Biodiversity of the southeastern United States: Upland terrestrial communities. John Wiley and Sons, New York.

Ecological Systems Placement

Ecological Systems Placement
Ecological System Unique ID Ecological System Name
CES202.329 Southern Piedmont Granite Flatrock and Outcrop


NatureServe Conservation Status
Global Status: G1 (14Dec1998)
Rounded Global Status: G1 - Critically Imperiled
Reasons: This community is limited to granite flatrocks in Georgia, extreme eastern Alabama, and extreme southwestern South Carolina. It occupies solution pools in flat exposures of granite and is characterized by a set of endemic plants. These pools do not occur on all flatrocks in the region and are vulnerable to degradation and destruction by recreational off-road vehicle traffic. Total acreage of this community is less 20 hectares (less than 50 acres).

Distribution
Color legend for Distribution Map
United States Distribution: AL, GA, SC
Global Distribution: United States
Global Range: This community is limited to granitic flatrocks of Georgia, South Carolina, and Alabama.

U.S. Forest Service Ecoregions
Domain Name: Humid Temperate Domain
Division Name: Subtropical Division
Province Name: Southeastern Mixed Forest Province
Province Code: 231 Occurrence Status: Confident or certain
Section Name: Southern Appalachian Piedmont Section
Section Code: 231A Occurrence Status: Confident or certain


Vegetation

Vegetation Summary: The vegetation of these pools is limited to the rare aquatic endemics Amphianthus pusillus, Isoetes melanospora, and Isoetes tegetiformans (Wharton 1978). All of these species are endemic to granitic flatrocks of the southeastern United States. Diamorpha smallii may occupy marginal areas of this community, which are flooded for shorter periods of time. Other species which may occur are Juncus georgianus and Cyperus granitophilus.

Vegetation Composition (incomplete)
Species Name Rounded Global Status Growth Form Stratum Charact-
eristic
Dominant Constant
Cover Class %
Con-
stancy
%
Amphianthus pusillus G1 Flowering forb Herb (field)  
 
 
Isoetes melanospora G1 Fern (Spore-bearing forb) Herb (field)  
 
 
Isoetes piedmontana G1 Fern (Spore-bearing forb) Herb (field)      
 
 
Isoetes tegetiformans G1 Fern (Spore-bearing forb) Herb (field)  
 
 
Cyperus granitophilus G1 Graminoid Herb (field)      
 
 


At-Risk Species Reported for this Association
Scientific Name
  (Common Name)
NatureServe Global Status U.S. Endangered Species Act Status
Amphianthus pusillus
  (Little Amphianthus)
G2 LT: Listed threatened
Cyperus granitophilus
  (Granite-loving Flatsedge)
G3G4Q  
Isoetes melanospora
  (Black-spored Quillwort)
G1 LE: Listed endangered
Isoetes tegetiformans
  (Merlin's-grass)
G1 LE: Listed endangered


Environmental Setting

Wetland Indicator: Y
Environmental Summary: This community occupies the deepest and wettest pools on granitic flatrocks of Georgia, South Carolina, and Alabama. These pools have intact rims, shallow soils, and are of sufficient depth to retain water for several weeks after a rain (Baker 1956).


Dynamic Processes


Plot Sampling & Classification Analysis

Plots stored in VegBank


Authors/Contributors
Concept Author(s): A.S. Weakley
Element Description Edition Date: 01Jan1996
Element Description Author(s): A.S. Weakley
NatureServe Conservation Status Factors Edition Date: 14Dec1998
NatureServe Conservation Status Factors Author(s): A.S. Weakley

Ecological data developed by NatureServe and its network of natural heritage programs (see Local Programs) and other contributors and cooperators (see Sources).


References
  • Baker, W. B. 1956. Some interesting plants of the granite outcrops of Georgia. Georgia Mineral Newsletter 9(1):10-19.

  • Burbanck, M. P., and R. B. Platt. 1964. Granite outcrop communities of the Piedmont Plateau in Georgia. Ecology 45:292-306.

  • Coffey, J. C. 1964. A floristic study of the flat granitic outcrops of the lower Piedmont, South Carolina. M.S. thesis, University of South Carolina, Columbia.

  • Nelson, J. B. 1986. The natural communities of South Carolina: Initial classification and description. South Carolina Wildlife and Marine Resources Department, Division of Wildlife and Freshwater Fisheries, Columbia, SC. 55 pp.

  • Quarterman, E., M. P. Burbanck, and D. J. Shure. 1993. Rock outcrop communities: Limestone, sandstone, and granite. Pages 35-86 in: W. H. Martin, S. G. Boyce, and A. C. Echternacht, editors. Biodiversity of the southeastern United States: Upland terrestrial communities. John Wiley and Sons, New York.

  • Schotz, Al. Personal communication. Community Ecologist. Alabama Natural Heritage Program. Huntingdon College, Massey Hall, 1500 East Fairview Avenue, Montgomery, AL 36106-2148.

  • Southeastern Ecology Working Group of NatureServe. No date. International Ecological Classification Standard: International Vegetation Classification. Terrestrial Vegetation. NatureServe, Durham, NC.

  • Wharton, C. H. 1978. The natural environments of Georgia. Georgia Department of Natural Resources, Atlanta. 227 pp.


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